Among his Carolina discoveries were four species of frogs (among them the handsome green tree frog, Hyla cinerea), three of turtles, and one species of lizard, found during his stay in Charleston. He also collected three new species of fish in Charleston harbor. He was especially interested in invertebrates, and the names of fourteen new species of coelenterates, mollusks, worms, and crabs that he described from South Carolina are still valid today, among them the familiar fiddler crab (Uca pugilator) of coastal salt marshes.